A list in temporal order follows the Google Scholar and ORCID links, which may not find all of my publications. The list is up to date as of 14 April 2019.

Google scholar Proctor publications

ORCID Proctor publications


  1. Proctor, H. and G. Pritchard. 1989. Neglected predators: water mites (Acari: Parasitengona: Hydrachnellae) in freshwater communities. Journal of the North American Benthological Society 8: 100-111.
  2. Proctor, H.C. 1989. Occurrence of protandry and a female-biased sex-ratio in a sponge-associated water mite (Acari: Unionicolidae). Experimental and Applied Acarology 7: 289-297.
  3. Proctor, H.C. and G. Pritchard. 1990. Variability in the life history of Unionicola crassipes, a sponge-associated water mite.  Canadian Journal of  Zoology 68: 1227-1232.
  4. Proctor, H.C. and G. Pritchard. 1990. Prey detection by the water mite Unionicola crassipes (Acari: Unionicolidae). Freshwater Biology 23: 271-290.
  5. Proctor, H.C. 1991. Courtship in the water mite Neumania papillator: males capitalize on female adaptations for predation. Animal Behaviour 42:589-598.
  6. Proctor, H.C. 1991. The evolution of copulation in water mites: a comparative test for nonreversing characters. Evolution 45: 558-567.
  7. Johnson, D.L. and H.C. Proctor. 1991. Effects of phytoseiid predators on the sex ratio of the spider mite Panonychus ulmi. Canadian Journal of Zoology  69: 208-212.
  8. Proctor, H.C. 1992. Mating and spermatophore morphology of water mites (Acari: Parasitengona). Zoological Journal of the Linnean Society 106: 341-384.
  9. Proctor, H.C. 1992. Discord between field and laboratory sex ratios of the water mite Neumania papillator Marshall (Acari: Unionicolidae). Canadian Journal of Zoology 70: 2483-2486.
  10. Proctor, H.C. 1992. Effect of food deprivation on mate-searching and spermatophore production in male water mites (Acari: Unionicolidae). Functional Ecology 6: 661-665.
  11. Proctor, H.C. 1992. Cohabitation of six species of spiders in webs of Cyrtophora moluccensis (Araneae, Araneidae) in Moorea, French Polynesia. Journal of Arachnology 20: 144-145.
  12. Proctor, H.C. 1992. Sensory exploitation and the evolution of male mating behaviour: a cladistic test using water mites (Acari: Parasitengona). Animal Behaviour 44: 745-752.
  13. Baker, R.L., M.R.L. Forbes and H.C. Proctor. 1992. Sexual differences in development and behaviour of larval Ischnura verticalis (Odonata: Coenagrionidae). Canadian Journal of Zoology 70: 1161-1165.
  14. Proctor, H.C. 1993. Mating biology resolves trichotomy for cheliferoid pseudoscorpions (Pseudoscorpionida, Cheliferoidea). The Journal of Arachnology 21: 156-158.
  15. Proctor, H.C. and L.D. Harder. 1994. Pollen load, capsule weight and seed production in three orchid species. Canadian Journal of Botany. 72: 249-255.
  16. Proctor, H.C. and B.P. Smith. 1994. Mating behaviour of the water mite Arrenurus manubriator (Acari: Arrenuridae). Journal of Zoology, London 232: 473-483.
  17. Proctor, H.C., R.L. Baker, and D.T. Gwynne. 1995. Mating behaviour and spermatophore morphology: a comparative test of the female choice hypothesis. Canadian Journal of Zoology, 73: 2010-2020.
  18. Proctor, H.C. and L.D. Harder. 1995. Effect of pollination success on floral longevity in the orchid Calypso bulbosa (Orchidaceae). American Journal of Botany, 82: 1131-1136.
  19. Proctor, H.C. 1997. Mating behavior of Physolimnesia australis (Halik) (Acari: Limnesiidae), a non-parasitic, rotifer-eating water mite from Australia. Journal of Arachnology 25: 321-325.
  20. Proctor, H.C., H.M. Gray and B.M. OConnor. 1997. Subaquatic mites (Acari: Astigmata) associated with adult freshwater leeches (Hirudinea: Erpobdellidae). Journal of Natural History 31: 539-544.
  21. Garga, N., H. Proctor and R. Belczewski. 1997. Leg size affects mating success in Tarsonemus confusus Ewing (Prostigmata: Tarsonemidae). Acarologia 38: 369-375.
  22. Walter, D.E. and H. C. Proctor. 1998. Feeding behaviour and phylogeny: observations on early derivative Acari. Experimental and Applied Acarology 22: 39-50.
  23. Proctor, H.C. 1998. Indirect sperm transfer in arthropods. Annual Review of Entomology 43: 153-174.
  24. Proctor, H.C. 1998. Effect of pollen age on fruit set, fruit weight, and seed set in three orchid species. Canadian Journal of Botany 76: 420-427.
  25. Walter, D.E. and H.C. Proctor. 1998. Predatory mites in tropical Australia: local species richness and complementarity. Biotropica 30: 72-81.
  26. Proctor, H.C. 1999. Gallilichus jonesi sp. n. (Acari: Ascouracaridae), a new species of feather mite from the quills of the Australian Brush-Turkey (Aves: Megapodiidae). Australian Journal of Entomology, 38: 77-84.
  27. Proctor, H.C. and I. Owens. 2000. Mites and birds: diversity, parasitism and coevolution. Trends in Ecology and Evolution 15: 358-364.
  28. Proctor, H.C. 2001. Extracting aquatic mites from stream substrates: a comparison of three methods. Experimental and Applied Acarology 25: 1-11.
  29. Proctor, H.C. 2001. Megninia casuaricola sp. n. (Acari: Analgidae), the first feather mite from a cassowary (Aves: Struthioniformes: Casuariidae). Australian Journal of Entomology 40: 335-341.
  30. Williams, C.R. and H.C. Proctor. 2002. Parasitism of mosquitoes (Diptera: Culicidae) by larval mites (Acari: Parasitengona) in Adelaide, South Australia. Australian Journal of Entomology 41: 161-163.
  31. Göth, A and H. Proctor. 2002. Pecking preferences in hatchlings of the Australian brush-turkey, Alectura lathami (Megapodiidae): the role of food type and colour. Australian Journal of Zoology 50: 93-102.
  32. Proctor, H., K. Montgomery, K. Rosen and R.L. Kitching. 2002. Are tree trunks habitats or highways? A comparison of oribatid mite assemblages from hoop-pine bark and litter. Australian Journal of Entomology 41: 294-299.
  33. Proctor, H.C. 2003. Feather mites (Acari: Astigmata): ecology, behavior and evolution. Annual Review of Entomology 48: 185-209.
  34. Nakamura, A., H. Proctor and C.P. Catterall. 2003. Using soil and litter arthropods to assess the state of rainforest restoration. Ecological Management & Restoration 4 (supplement): S20-S28.
  35. Mironov, S.V., J. Dabert and H. Proctor. 2003. New feather mites of the family Pterolichidae (Acari: Pterolichoidea) from parrots (Aves: Psittaciformes) in Australia. Australian Journal of Entomology 42: 185-202.
  36. Fan Q.-H., Walter D.E. and Proctor H.C. 2003. A review of the family Barbutiidae (Acari: Raphignathoidea), with the description of two new species from Australia. Systematic & Applied Acarology 8: 107-130.
  37. Kanowski, J., Catterall, C. P., Wardell-Johnson, G. W., Proctor, H. and Reis, T. 2003. Development of forest structure on cleared rainforest land in eastern Australia under different styles of reforestation. Forest Ecology and Management 183: 265-280.
  38. Fan, Q.-H., Walter, D.E., and Proctor, H. 2003. A review of the genus Ledermuelleriopsis Willmann (Acari: Prostigmata: Stigmaeidae). Invertebrate Systematics 17: 551-574.
  39. Proctor, H.C., J. Kanowski, G. Wardell-Johnson, T. Reis and C.P. Catterall. 2003. Does diversity beget diversity? A comparison between plant and leaf-litter invertebrate richness from pasture to rainforest. Records of the South Australian Museum Monograph Series No. 7: 267-274.
  40. Proctor, H.C. and Jones, D.N. 2004. Geographical structuring of feather mite assemblages from the Australian brush-turkey (Aves: Megapodiidae). Journal of Parasitology 90: 60-66.
  41. Boulton, A., M. Harvey and H. Proctor. 2004. Of spates and species: responses by interstitial water mites to simulated spates in a subtropical Australian river. Experimental and Applied Acarology 34: 149-169.
  42. Proctor, H.C. and N. Garga. 2004. Red, distasteful water mites: did fish make them that way? Experimental and Applied Acarology 34: 127-147.
  43. Lian Pin Koh, R.R. Dunn, N.S. Sodhi, R.K. Colwell, H.C. Proctor, and V.S. Smith. 2004. Species coextinctions and the biodiversity crisis.  Science 305: 1632-1634.
  44. Mironov, S.V. and H.C. Proctor. 2005. A new feather mite genus of the family Psoroptoididae (Acari: Analgoidea) from cassowaries.  Journal of Natural History 39: 3297-3304.
  45. Bochkov, A.V., G. Williams and H. Proctor. 2005. First record of Picobia zumpti (Acari: Syringophilidae) from quills of the rock pigeon in North America and description of the male.  Belgian Journal of Entomology 7: 121-127.
  46. Proctor, H and A. Grigg. 2006. Aquatic invertebrates in final void water bodies at an open-cut coal mine in central Queensland.  Australian Journal of Entomology 45: 107-121.
  47. Dabert, J., S.V. Mironov and H. Proctor. 2006. A new species of the genus Titanolichus Gaud & Atyeo, 1996 (Acari: Astigmata: Pterolichidae) from the endangered orange-bellied parrot Neophema chrysogaster (Aves: Psittaciformes) from Australia. Australian Journal of Entomology 45: 206-214.
  48. Beaulieu, F., D. E. Walter, H. C. Proctor, R. L. Kitching and F. Menzel. 2006. Mesostigmatid mites (Acari: Mesostigmata) on rainforest tree trunks: arboreal specialists, but substrate generalists?  Experimental and Applied Acarology 39: 25-40.
  49. Knee, W. and H. Proctor. 2006. Keys to the families and genera of blood and tissue feeding mites associated with Albertan birds.  Canadian Journal of Arthropod Identification 2: 1-18.
  50. Proctor, H., G. Zimmerman and K. Meyer. 2006. A new feather mite, Aetacarus elanoides sp. n. (Acari: Gabuciniidae), from the Swallow-tailed Kite Elanoides forficatus (Linnaeus) (Falconiformes: Accipitridae; Perninae). Zootaxa 1252: 37-47.
  51. Grimbacher, P.S., C.P. Catterall, J. Kanowski and H.C. Proctor. 2007. Responses of ground-active beetle assemblages to different styles of reforestation on cleared rainforest land. Biodiversity and Conservation 16: 2167-2184.
  52. Boeckner, M. and H. Proctor. 2005*. Water-bears from the Rocky Mountains: a first look at Alberta’s tardigrade fauna. Canadian Field Naturalist 119: 586-588 [*note: although issue is dated 2005, it was not actually published until May 2007]
  53. Knee, W. and H. Proctor. 2007. Host records for the northern fowl mite, Ornithonyssus sylviarum (Mesostigmata: Macronyssidae), from birds of North America (Canada, USA, and Mexico). Journal of Medical Entomology 44: 709-713.
  54. Mironov, S.V., H.C. Proctor, M. Barreto and G. Zimmerman. 2007. New genera and species of feather mites of the family Gabuciniidae (Astigmata: Pterolichoidea) from New World raptors (Aves: Falconiformes).  The Canadian Entomologist 139: 757-777.
  55. Kelly, J.R., H. Proctor and J.P. Volpe. 2008. Intertidal community structure differs significantly between substrates dominated by native eelgrass (Zostera marina L.) and adjacent to the introduced oyster Crassostrea gigas (Thunberg) in British Columbia, Canada. Hydrobiologia 596: 57-66.
  56. Gentes, M.-L., H. Proctor and G. Wobeser. 2007. Demodicosis in a mule deer (Odocoileus hemionus hemionus) from Saskatchewan, Canada. Journal of Wildlife Diseases 43: 758–761.
  57. Galván, I., E. Barba, R. Piculo, J. L. Cantó, V. Cortés, J. S. Monrós, F. Atiénzar and H. Proctor. 2008. Feather mites and birds: an interaction mediated by uropygial gland size? Journal of Evolutionary Biology 21: 133-144.
  58. Knee, W., H. Proctor and T. Galloway. 2008. Survey of nasal mites (Rhinonyssidae, Ereynetidae, and Turbinoptidae) associated with birds in Alberta and Manitoba, Canada. The Canadian Entomologist 140: 364-379.
  59. Mironov, S.V. and H.C. Proctor. 2008. The probable association of feather mites of the genus Ingrassia (Analgoidea: Xolalgidae) with the blue penguin Eudyptula minor (Aves: Spenisciformes) in Australia. Journal of Parasitology 94: 1243-1248.
  60. Piper, S.D., C.P. Catterall, J.J. Kanowski and H.C. Proctor. 2009. Biodiversity recovery during rainforest reforestation as indicated by rapid assessment of epigaeic ants in tropical and subtropical Australia. Austral Ecology 34: 422-434.
  61. Williams, B.W., S.R. Gelder and H. Proctor. 2009. Distribution and first reports of Branchiobdellida (Annelida: Clitellata) on crayfish in the Prairie Provinces of Canada. Western North American Field Naturalist 69: 119-124.
  62. Boeckner, M.J., J. Sharma and H.C. Proctor. 2009. Revisiting the meiofauna paradox: dispersal and colonization of nematodes and other meiofaunal organisms in low- and high-energy environments. Hydrobiologia: 624: 91–106.
  63. Proctor, H. 2009. Can freshwater mites act as forensic tools? Experimental and Applied Acarology 49: 161-165.
  64. Mironov, S. and H. Proctor. 2009. Feather mites of the genus Proterothrix Gaud (Astigmata: Proctophyllodidae) from parrotbills (Passeriformes: Paradoxornithidae) in China. Journal of Parasitology: 95: 1093-1107.
  65. Stephenson, S., S. Hannon and H. Proctor. 2009. The function of feathers in tree swallow nests: insulation or ectoparasite barrier? The Condor 111: 479-487.
  66. Mironov, S. and H. Proctor. 2009. Atelophyllodes gen. n., a new feather mite genus of the family Proctophyllodidae (Astigmata: Analgoidea) from lyrebirds (Passeriformes: Menuridae). Zootaxa 2326: 51–61.
  67. Knee, W. and H. Proctor. 2010. Interactive HTML-based dichotomous key to female Rhinonyssidae (Mesostigmata) from birds in Canada. Canadian Journal of Arthropod Identification 9 (doi: 10.3752/cjai.2010.09: available online at
  68. Walter, D.E. and H.C. Proctor. 2010. Mites as modern models: acarology in the 21st century. Acarologia 50: 131-141 (invited paper for Acarologia’s 50th anniversary).
  69. Beaulieu, F., D.E. Walter, H.C. Proctor and R.L. Kitching. 2010. The canopy starts at 0.5 m: predatory mites (Acari: Mesostigmata) differ between rain forest floor soil and suspended soil at any height. Biotropica 42: 704-709.
  70. Kontschán, J., H. Proctor and J. Newton. 2010. Trachyuropoda kinsella n. sp. (Acari: Uropodina: Trachyuropodidae) from Alberta, Canada, with a key to Trachyuropoda species from temperate North America. International Journal of Acarology 36: 211-220.
  71. Carleton, R.E. and H.C. Proctor. 2010. Feather mites associated with Eastern Bluebirds (Sialia sialis L.) in Georgia, including the description of a new species of Trouessartia (Analgoidea: Trouessartiidae). Southeastern Naturalist 9: 605-623.
  72. Haribal, M. , H. Proctor, A.A. Dhondt, and E. Rodriguez. 2011. Biology of House Finch feather mites, Proctophyllodes pinnatus (Acari: Proctophyllodidae), parallels variation in preen gland secretions. International Journal of Acarology 37: 75 – 90.
  73. Mori, B.A., H.C. Proctor, D.E. Walter and M.L. Evenden. 2011. Phoretic mite associates of mountain pine beetle at the leading edge of an infestation in northwestern Alberta, Canada. The Canadian Entomologist 143: 44-55.
  74. Mestre, A., F. Mesquita-Joanes, H. Proctor and J. S. Monrós. 2011. Different scales of spatial segregation of two species of feather mites on the wings of a passerine bird. Journal of Parasitology 97: 237-244.
  75. Gale, K.S.P. and H.C. Proctor. 2011. Diets of two congeneric species of crayfish worm (Annelida: Clitellata; Branchiobdellidae) from western Canada. Canadian Journal of Zoology 89: 289-296.
  76. Williams, B.W., H.C. Proctor and T. Clayton. 2011. Range extension of the Northern Crayfish, Orconectes virilis (Decapoda, Cambaridae), in the western prairie provinces of Canada. Crustaceana 84: 451-460.
  77. Valim, M.P., F.A. Hernandes and H.C. Proctor. 2011. Feather mites of Brazil (Acari: Astigmata: Analgoidea and Pterolichoidea). International Journal of Acarology 37: 293-324.
  78. Williams, B.W., K.L. Williams, S.R. Gelder and H.C. Proctor. 2011. Distribution of Entocytheridae (Crustacea: Ostracoda) in the northern prairies of North America and reports of opportunistic clitellate annelids on crayfish hosts. Western North American Naturalist 71: 276-282.
  79. Mironov, S.V. and H.C. Proctor. 2011. Four new feather mite species of the family Pteronyssidae (Astigmata: Analgoidea) from laughing-thrushes (Passeriformes: Timaliidae) in China. Acarina 19: 35-51.
  80. Newton, J.S., J. Glasier, H.E.L. Maw, H.C. Proctor and R.G. Foottit. 2011. Ants and subterranean Sternorrhyncha in a native grassland in east-central Alberta, Canada. The Canadian Entomologist 143: 518-523.
  81. Locke, S.A, A.R. Lapierre, K. Byers, H. Proctor, J.D. McLaughlin and D.J. Marcogliese. 2012. Molecular and morphological evidence for the Holarctic distribution of Urogonimus macrostomus (Rudolphi, 1803) Monticelli, 1888 (Digenea: Leucochloridiidae). Journal of Parasitology 98: 880-882.
  82. Williams, B.W., Stuart R. Gelder, Heather, C. Proctor and David W. Coltman. 2012. Molecular phylogeny of North American Branchiobdellida (Annelida: Clitellata). Molecular Phylogenetics & Evolution 66: 30-42.
  83. Barreto, M. M. E. Burbano, H. C. Proctor, S. V. Mironov and G. Wauthy. 2012. Feather mites (Acari) from Colombia. Zootaxa 3516: 1–68.
  84. Cameron, E.K., K.M. Knysh, H.C. Proctor and E.M. Bayne. 2013. Influence of two exotic earthworm species with different foraging strategies on abundance and composition of boreal microarthropods. Soil Biology and Biochemistry 57: 334-340 (accepted 17 July 2012).
  85. Newton, J.S. and H.C. Proctor. 2013. A fresh look at weight-estimation models for soil mites (Acari). International Journal of Acarology 39: 72-85, DOI:10.1080/01647954.2012.744351 (accepted 23 October 2012).
  86. Cameron E.K., H.C. Proctor and E.M. Bayne. 2013. Effects of an ecosystem engineer on belowground movement of microarthropods. PLoS ONE 8(4): 9 pp. e62796, doi:10.1371/journal.pone.0062796 (accepted 25 March 2013).
  87. Byers, K. and H. Proctor. 2013. Symbiotic arthropods from the house sparrow (Passer domesticus) from two locations in Alberta, Canada. The Canadian Entomologist 145: 668-673.
  88. Diaz-Aguilar, I. S.A. Quideau, H.C. Proctor , B.E. Kishchuk and J.R. Spence.2013. Influence of stand composition on predatory mite (Mesostigmata) assemblages from the forest floor in western Canadian boreal mixedwood forests. Forest Ecology and Management 309: 105-114.
  89. Glasier, J.R.N., J.H. Acorn, S. Nielsen and H. Proctor. 2013. Ants (Hymenoptera: Formicidae) of Alberta: a key to species based primarily on the worker caste. Canadian Journal of Arthropod Identification 22 (, DOI: 10.3752/cjai.2013.22).
  90. Anholt, H., C. Himsworth, J. Rothenburger, H. Proctor, and D.M. Patrick. 2014. Ear mange mites (Notoedres muris) in black and Norway rats (Rattus rattus and Rattus norvegicus) from inner-city Vancouver, Canada. Journal of Wildlife Diseases 50: 104-108. DOI: 10.7589/2013-02-046
  91. Byers, K.A. and H.C. Proctor. 2014. Like a glove: do the dimensions of male adanal suckers and tritonymphal female docking papillae correlate in the Proctophyllodidae (Astigmata: Analgoidea)? Acarologia 54: 3-14. DOI: 10.1051/acarologia/20142110
  92. Fernández-González, S., A. Pérez-Rodríguez, I. de la Hera, H. C. Proctor and J. Pérez-Tris. 2015. Different space preferences and within-host competition promote niche partitioning between symbiotic feather mite species. International Journal for Parasitology 45: 655-662
  93. Wang, Z.-Y. and H. Proctor. 2015. Two new feather mites of the genus Neocalcealges Orwig (Analgoidea: Trouessartiidae) from the Sichuan province of China. Zootaxa 3946: 567–576 DOI:
  94. Stefan, L.M., E. Gómez-Díaz, E. Elguero , H.C. Proctor, K.D. McCoy and J. González-Solís. 2015. Niche partitioning of feather mites within a seabird host, Calonectris borealis. PLoS ONE 10(12): e0144728. doi:10.1371/journal.pone.0144728
  95. Dabert, M., H. Proctor and J. Dabert. 2016. Higher-level molecular phylogeny of the water mites (Acariformes: Prostigmata: Parasitengonina: Hydrachnidiae). Molecular Phylogenetics and Evolution 101: 75-90.
  96. Doña,J., H. Proctor, S. Mironov, D. Serrano and R. Jovani. 2016. Global associations between birds and vane-dwelling feather mites. Ecology 97: 3242. DOI: 10.1002/ecy.1528.
  97. Jiang, X., Z. Song, J. Xiong, H. Proctor and Z. Xie. 2017. Different surrogacy approaches for stream macroinvertebrates in discriminating human disturbances in Central China. Ecological Indicators 75: 182-191.
  98. Kiedrowicz, A., L. Kuczyński, M. Lewandowski, H. Proctor and A. Skoracka. 2017. Behavioural responses to potential dispersal cues in two economically important species of cereal-feeding eriophyid mites. Scientific Reports 7: 3890. DOI:
  99. Haché, S., E. Bayne, M.-A. Villard, H. Proctor, C. Davis, D. Stralberg, J. Janes, M. Hallworth, K. Foster, E. Chidambara-Vasi, A. Grossi, J. Gorrell, and R. Krikun.2017. Phylogeography of a migratory songbird across its Canadian breeding range: implications for conservation units. Ecology and Evolution 7: .
  100. Carlson, C.J. K.R. Burgio, E.R. Dougherty, A.J. Phillips, V.M. Bueno, C.F. Clements, G. Castaldo, T.A. Dallas, C.A. Cizauskas, G.S. Cumming, J. Doña, N.C. Harris, R. Jovani, S. Mironov, O.C. Muellerklein, H.C. Proctor, W.M. Getz. 2017. Parasite biodiversity faces extinction and redistribution in a changing climate. Science Advances 3: e1602422.
  101. Doña J, H. Proctor, S. Mironov , D. Serrano and R. Jovani. 2017. Host specificity, infrequent major host switching and the diversification of highly host-specific symbionts: the case of vane-dwelling feather mites. Global Ecology and Biogeography 2017;00:1–11.
  102. De Castro, L.A.S., C.E. Da Rocha and H.C. Proctor. 2017. Morphological studies on the water mite Hydryphantes ramosus Daday, 1905 (Acariformes: Hydrachnidiae: Hydryphantidae) from Brazil, with a description of the larva. Systematic and Applied Acarology, 22:1688-1697
  103. Fernández-González S, Pérez-Rodríguez A, Proctor HC, De la Hera I, Pérez-Tris J. 2018. High diversity and low genetic structure of feather mites associated with a phenotypically variable bird host. Parasitology (published online: 17 January 2018, pp. 1-8).
  104. Matthews, A.E., P.B. Klimov, H.C. Proctor, A.P.G. Dowling, L. Diener, S.B. Hager, J.L. Larkin, D.W. Raybuck, C.J. Fiss, D.J. McNeil and T.J. Boves. 2018. Cophylogenetic assessment of New World warblers (Parulidae) and their symbiotic feather mites (Proctophyllodidae). Journal of Avian Biology 2018: e01580, doi:10.1111/jav.01580
  105. Proctor, H. and W. Knee. 2018. Asymmetry and polymorphism in males of the feather mite Michaelia neotropica Hernandes and Mironov (Acariformes: Astigmata: Freyanidae). Zoologisher Anzeiger 273: 226-230
  106. Doña J., H. Proctor H, D. Serrano D, K.P. Johnson, A. Oddy-van Oploo, J.C. Huguet-Tapia, M.S. Ascunce and R. Jovani. 2018. Feather mites play a role in cleaning host feathers: new insights from DNA metabarcoding and microscopy. Molecular Ecology, 00:1–16,
  107. Goater, C.P.J. Dyck, H. Proctor and K. Floate. 2018. Hyperparasitism of an avian ectoparasitic hippoboscid fly, Ornithomya anchineuria, by the mite, Myialges cf. borealis, in Alberta, Canada. Journal of Parasitology 104: 111-116.
  108. Vanstreels, R.E.T., H. Proctor, A. Snyman, R. Hurtado, K. Ludynia, N.J. Parsons and P.A. Pistorius. 2018. Nasal mites (Mesostigmata: Rhinonyssidae) in African penguins (Spheniscus demersus). Parasitology 1–7.
  109. Meehan, M.L, Z. Song and H. Proctor. 2018. Roles of environmental and spatial factors in structuring assemblages of forest-floor Mesostigmata in the boreal region of Northern Alberta, Canada. International Journal of Acarology 44: 300-309, DOI:10.1080/01647954.2018.1520297.
  110. Beaulieu F., Knee W., Nowell V., Schwarzfeld M., Lindo Z., Behan‑Pelletier V.M., Lumley L., Young M.R., Smith I., Proctor H.C., Mironov S.V., Galloway T.D., Walter D.E., Lindquist E.E. (2019) Acari of Canada. In: Langor DW, Sheffield CS (Eds) The Biota of Canada – A Biodiversity Assessment. Part 1: The Terrestrial Arthropods. ZooKeys 819: 77–168.
  111. Meehan, M.L., Z. Song, L.M. Lumley, T.P. Cobb and H. Proctor. 2019. Soil mites as bioindicators of disturbance in the boreal forest in northern Alberta, Canada: testing taxonomic sufficiency at multiple taxonomic levels. Ecological Indicators 102: 349-365.
  112. Wijerathna, A., C. Whitehouse, H. Proctor and M. Evenden. 2019. Testing for trade-offs between flight and reproduction in the mountain pine beetle (Coleoptera: Curculionidae) on two pine (Pinaceae) hosts. The Canadian Entomologist, 151: 298-310.
  113. Martens, A.A. U. Silins, H. C. Proctor, C. H. S. Williams, M. J. Wagner, M.B. Emelko and M. Stone. 2019. Long-term impact of severe wildfire and post-wildfire salvage logging on macroinvertebrate assemblage structure in Alberta’s Rocky Mountains. International Journal of Wildland Fire, 28:738-749.
  114. Durkin, E.S., H. Proctor and L. Luong. 2019. Life history of Macrocheles muscaedomesticae (Parasitiformes: Macrochelidae): new insights on life history and evidence of facultative parasitism on Drosophila. Experimental and Applied Acarology, 79: 309–321.
  115. Young, M.R., H. Proctor, J.R. DeWaard and P.D.N. Hebert. 2019. DNA barcodes expose unexpected diversity in Canadian mites. Molecular Ecology, 28:5347–5359.
  116. Wiecek, M., W. Szydlo, Wiktoria, J. Dabert and H. Proctor. 2019. Delimiting species of water mites of the genus Hydrodroma (Acari: Hydrachnidiae: Hydrodromidae) from North America and Europe: Integrative evidence of species status from COI sequences and morphology. Zoologischer Anzeiger, 284:16–29.
  117. Grossi, A.A. and H. Proctor. 2020. The distribution of quill mites (Betasyringophiloidus seiuri) among flight feathers of the Ovenbird (Seiurus aurocapilla). Journal of Parasitology, 106:82-89.
  118. Song, Z. and H. Proctor. 2020. Parasite prevalence in intermediate hosts increases with waterbody age and abundance of final hosts. Oecologia, 192:311–321.
  119. Byers, K.A. and H. Proctor. 2020. Morphology of genitalia and non-genitalic contact structures in Trouessartia feather mites (Astigmata: Analgoidea: Trouessartiidae): is there evidence of correlated evolution between the sexes? Canadian Journal of Zoology, 98:815–826.
  120. Worthington, A.A., C.D. Kelly and H. Proctor. 2020. Weak relationships of parasite infection with sexual and life-history traits in wild-caught Texas field crickets (Gryllus texensis). Ecological Entomology, 46:76–88.
  121. Song, Z. and H. Proctor. 2020. Application of DNA barcoding and morphometric analysis in differentiating cystacanths of Polymorphus species (Acanthocephala: Polymorphidae) from central Alberta, Canada. Parasitology Research, 119:3359-3368. doi: 10.1007/s00436-020-06870-6
  122. de Castro, L.A.S., H. Proctor and A.C. Lofego. 2020. A new species of Clathrosperchonella Lundblad 1937 (Acariformes: Hydrachnidiae: Hydryphantoidea: Rhynchohydracaridae) from Brazil, with descriptions of the female, male and larva. Systematic and Applied Acarology, 25:1745-1753.
  123. Grossi, A.A. and H. Proctor. 2021. Variation in ectosymbiont assemblages associated with Rock Pigeons (Columba livia) from coast to coast in Canada. Diversity, 13:1-18.


  1. Walter, D.E. and H.C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. University of New South Wales Press and CAB International.
  2. Halliday, R. B., Walter, D. E., Proctor, H. C., Norton, R. A. and Colloff, M. J. (eds.) 2001. Acarology: Proceedings of the 10th International Congress. CSIRO Publishing, Melbourne.
  3. Proctor, H.C. (ed.) 2004. Aquatic Mites: From Genes to Communities. Kluwer Academic Publishers, Dordrecht (hardcopy book version of Experimental & Applied Acarology 34, Nos 1-2, 2004).
  4. de Moraes, G. J. and H. Proctor (eds.) 2011. Acarology XIII: Proceedings of the International Congress. Zoosymposia 6: 1-304. Magnolia Press, Auckland (note: all open access
  5. Walter, D.E. and H.C. Proctor. 2013. Mites – Ecology, Evolution and Behaviour (2nd edition): Life at a Microscale. Springer. 486 pp. ISBN 978-94-007-7163-5.


  1. Proctor, H.C. 1996. Behavioral characters and homoplasy: perception versus practice. pp. 131-149 in Homoplasy: the Recurrence of Similarity in Evolution, edited by M.J. Sanderson and L. Hufford. Academic Press. (refereed)
  2. Proctor, H.C. 1999. Spermatophores in the Arthropods. pp. 578-586 in The Encyclopedia of Reproduction, Vol. 4., edited by E. Knobil and J.D. Neill. Academic Press.
  3. Proctor, H.C. 2003. Male mating effort and female remating in the water mite Arrenurus manubriator (Acari: Arrenuridae). pp. 211-221 in I.M. Smith (ed.) An Acarological Tribute to Dave Cook. Indira Publishing House, West Bloomfield.  ISBN 0-930337-18-2 (refereed).
  4. Catterall, C.P., Kanowski, J., Wardell-Johnson, G., Proctor, H.C., Reis, T., Harrison, D.A., & Tucker, N.I.J. 2004. Quantifying the biodiversity values of reforestation: perspectives, design issues and outcomes in Australian rainforest landscapes. Lunney, D. (ed.) Conservation of Australia’s Forest Fauna. 2nd edition Royal Zoological Society of New South Wales, Mosman, NSW, Australia. Pp. 359-393.
  5. Kanowski, J., C.P. Catterall, H. Proctor, T. Reis, N.I.J. Tucker & G. Wardell-Johnson. 2005. Chapter 12. Biodiversity values of timber plantations and restoration plantings for rainforest fauna in tropical and sub-tropical Australia. Pp. 183-205. In: ‘Reforestation in the tropics and sub-tropics of Australia using rainforest tree species’ (Eds Erskine, P.D., Lamb, D. and Bristow, M.). Rural Industries Research and Development Corporation, Canberra, and the Rainforest CRC, Cairns.
  6. Proctor, H.C. 2009. Hydrachnida (water mites). Chapter 176 in Encyclopedia of Inland Waters (G. Likens, ed.). Volume 2. pp. 335-345. Elsevier Ltd., Oxford. (refereed)
  7. Valdecasas, A.G., M. Aboal, S. Cirujano, S. Iepure, D. Jaume, H. Proctor and J.L. Velasco. 2010. Sampling continental freshwaters. Chapter 10 (pp. 213-272) in ABC Taxa: Volume 8 – Manual on Field Recording Techniques and Protocols for All Taxa Biodiversity Inventories. Editors: Jutta Eymann, Jérôme Degreef, Christoph Häuser, Juan Carlos Monje, Yves Samyn & Didier VandenSpiegel. date of pub: 2010. URL:
  8. Terry D. Galloway, Heather Proctor, and Sergei V Mironov. 2014. Chewing lice (Insecta: Phthiraptera: Amblycera, Ischnocera) and feather mites (Acari: Astigmatina: Analgoidea, Pterolichoidea): ectosymbionts of grassland birds in Canada. Pages: 141-190. Eds: H. A. Cárcamo and D. J. Giberson. Arthropods of Canadian Grasslands (Volume 3). Publisher: Biological Survey of Canada. DOI: doi: http://dx.10.3752/9780968932162.ch5. Available for download at
  9. Proctor, H.C., Smith, I.M., Cook, D.R., Smith, B.P. 2015. Subphylum Chelicerata, Class Arachnida. In: Thorp, J., Rogers, D.C. (Eds.), Ecology and General Biology: Thorp and Covich’s Freshwater Invertebrates, Academic Press, 599–660.


  1. Proctor, H.C. 1996. Sex ratios and chromosomes in water mites (Acari: Parasitengona). pp. 441-445 in Acarology IX (Volume 1), Proceedings of the 9th International Congress of Acarology, edited by R. Mitchell, D.J. Horn, G.R. Needham, W.C. Welbourn. Ohio Biological Survey, Ohio.
  2. Proctor, H.C. and K. Wilkinson. 2001. Coercion and deceit: water mites (Acari: Hydracarina) and the study of intersexual conflict. pp. 155-169 In Halliday, R. B., Walter, D. E., Proctor, H. C., Norton, R. A. and Colloff, M. J. (eds.) Acarology: Proceedings of the 10th International Congress. CSIRO Publishing, Melbourne.
  3. Proctor, H.C. 2007. Aquatic mites in assessments of stream invertebrate diversity. pp. 105-117 in Acarology XI: Proceedings of the International Congress. Morales-Malacara, J.B., Behan-Pelletier, V., Ueckermann, E., Pérez, T.M., Estrada-Venegas, E.G. and Badii, M. (Eds.). Instituto de Biololgía; Facultad de Ciencias; Universidad Nacional Autónoma de México; Sociedad Latinoamericana de Acarología. México, 2002.
  4. Walter, D.E. and H.C. Proctor. 2007. Using your genetic system to win friends and obfuscate theorists: patterns in the distribution of genetic sex determination in the Parasitiformes. pp. 637-643 in Acarology XI: Proceedings of the International Congress. Morales-Malacara, J.B., Behan-Pelletier, V., Ueckermann, E., Pérez, T.M., Estrada-Venegas, E.G. and Badii, M. (Eds.). Instituto de Biololgía; Facultad de Ciencias; Universidad Nacional Autónoma de México; Sociedad Latinoamericana de Acarología. México, 2002.
  5. Proctor, H.C., G. Williams and D.H. Clayton. 2009. Population density and male polymorphism in the feather mite Falculifer rostratus (Acari: Falculiferidae). pp. 299-302 in M.W. Sabelis & J. Bruin (eds.) Trends in Acarology – Proceedings of the 12th International Congress. Springer [note: book is due Jan 2011, but editors say publication date for chapters is 2009]
  6. Proctor, H., J. Kanowski, C.P. Catterall, G. Wardell-Johnson and T. Reis. 2011. Rainforest-restoration success as judged by assemblages of soil- and litter-dwelling mites (Arachnida: Acari). pp. 234 – 254 in Acarology XIII: Proceedings of the International Congress. Zoosymposia 6: 1-304. de Moraes, G. and H. Proctor (editors)


  1. Proctor, H.C. and A. Grigg. 1999. An investigation of aquatic invertebrates at Moura Mine.  Report to Moura Mine BHP Coal Pty Ltd.
  2. Greenway, M., C. Polson and H.C. Proctor. 2001. A comparison of aquatic biota and water quality in two tributaries of Native Dog Creek with different land-use activities. pp. 245-252 In Rutherfurd, I., Sheldon, F., Brierley, G. and Kenyon, C. (eds.) Third Australian Stream Management Conference. Published by the Cooperative Research Centre for Catchment Hydrology, Brisbane.
  3. Grigg, A. and H.C. Proctor. 2001. Investigations into litter decomposition on the tailings dam at Kidston Gold Mine.  Report prepared for Kidston Gold Mine by the Centre for Mined Land Rehabilitation, University of Queensland.
  4. Wardell-Johnson, G., Kanowski, J., Catterall, C., Proctor, H. and Reis, T. 2001. Measuring the restoration of rainforest biodiversity: a case study in research design, and its implications for establishing monitoring networks. pp. 72-81 in Boyes, B. (ed.) Biodiversity – The Big Picture. Proceedings of the 2001 Southern Queensland Biodiversity Recovery Conference.
  5. Proctor, H.C. 2005. Water quality & macroinvertebrate assemblages of water bodies on and near Grasslands National Park, Saskatchewan. Report prepared for Grasslands National Park, Saskatchewan. 13 May 2005. 66 pp.